A method for quantifying biodiversity loss and its application to a 50-year record of deforestation across Madagascar

1 Conservation Science Program, World Wildlife Fund–US, 1250 24th Street NW, Washington, DC 20010, USA 2 Department of Environmental Science, Policy and Management, 137 Mulford Hall, University of California, Berkeley, CA 94720-3114, USA (current address) 3 Department of Environment and Climate Change, PO Box 402 Armidale NSW 2350, Australia 4 CSIRO Entomology GPO Box 1700, Canberra, ACT 2601, Australia (current address) 5 Department of Entomology, California Academy of Sciences, Golden Gate Park, San Francisco, CA 94118, USA 6 Center for Applied Biodiversity Science, Conservation International, Arlington, VA 22202, USA 7 Department of Natural Resources & Environmental Sciences, University of Illinois, Urbana, IL 61801, USA 8 Département Systématique et Evolution, Muséum National d’Histoire Naturelle, Paris Cedex 05, France 9 Department of Entomology, Natural History Museum, London, SW7 5BD, UK 10 Carnegie Museum of Natural History, 4400 Forbes Avenue, Pittsburgh, PA, 15213, USA